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around the continents may have persisted and given rise to modern animals. However this may be, we must regard this deep zone as the birthplace of the fauna which has survived; as the ancestral home of all the improved metazoa.
The effect of life upon the bottom is more interesting than the place where it began, and we are now to consider its influence upon animals, all whose ancestors and competitors and enemies had previously been pelagic. The cold, dark, silent, quiet depths of the sea are monotonous compared with the land, but they introduced many new factors into the course of organic evolution.
It is doubtful whether the animals which first settled on the bottom secured any more food than floating ones, but they undoubtedly obtained it with less effort, and were able to devote their superfluous energy to growth and to multiplication, and thus to become larger and to increase in numbers faster than pelagic animals. Their sedentary life must have been favorable to both sexual and asexual multiplication, and the tendency to increase by budding must have been quickly rendered more active, and one of the first results of life on the bottom must have been to promote the tendency to form connected cormi, and to retain the connection between the parent and the bud until the latter was able to obtain its own food and to care for itself. The animals which first acquired the habit of resting on the bottom soon began to multiply faster than their swimming allies, and their asexually produced progeny, remaining for a longer time attached to and nourished by the parent stock, were much more favorably placed for rapid growth. As the animals of the bottom live on a surface, or at least a thin stratum, while swimming animals are distributed through solid space, the rapid multiplication of bottom animals must soon have led to crowding and to competition, and it quickly became harder and harder for new forms from the open water to force themselves among the old ones, and colonization soon came to an end.
The great antiquity of all the types of structure which are represented among modern animals is therefore what we should