Page:TheTreesOfGreatBritainAndIreland vol04B.djvu/373

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BETULA

Betula, Linnæus, Sp. Pl. 982 (1753); Bentham et Hooker, Gen. Pi. iii. 404 (1880); Winkler, in Engler, Pfanzenreich, iv. 61, Betulaceæ, 56 (1904).
Betulaster, Spach, in Ann. Sc. Nat. sér. 2, xv. 198 (1841).
Apterocaryon and Chamæbetula, Opiz, in Lotos, v. 258 (1855).

Deciduous trees or shrubs belonging to the order Betulaceæ. Bark smooth with longitudinal lenticels, often peeling off in papery strips, and becoming on old trunks thick and furrowed near the base. Branchlets of two kinds: long shoots with several leaves and axillary buds, no true terminal bud being formed; and short shoots or dwarf spurs, each with two (rarely one or three) leaves and a terminal bud. Buds viscid, elongated, ovoid, fully grown and green at midsummer, composed of imbricated scales, but with the two basal ones short and lateral, usually only four scales being visible externally ; inner scales accrescent, and marking in falling the base of the shoots with ring-like scars. Leaves alternate, simple, stalked, penninerved; serrate, dentate or incised. Stipules lateral, enclosing the leaf in the bud, fugacious. Flowers monececious, fertilised by the wind, in cylindrical catkins, composed of closely imbricated three-lobed scales, with three flowers on each scale. Male catkins,’ formed in the preceding autumn, clustered in the axils of the upper leaves of a long shoot, erect and naked during winter, pendulous in spring. Staminate flowers, with a one- to four- lobed calyx; stamens two, with short bifurcated filaments, each of the four branches bearing an erect half-anther, there being thus apparently twelve stamens on each scale. Pistillate catkins, solitary, or two to four in a raceme, terminal on the short shoots, and appearing with the leaves in spring. Pistillate flowers, without a calyx, two-celled, with one ovule in each cell; styles two, stigmatic at the apex. Cones, ripening *® usually in autumn, composed of woody three-lobed scales and small fruits, deciduous together; nutlets oval or obovate, compressed, bearing the persistent styles at the apex, and with the outer shell produced into a marginal transparent wing, interrupted at the apex; seed solitary, pendulous, without albumen.®

In winter, species of Betula are readily distinguished by the short shoots on the older wood, which end in a terminal bud, and are densely clothed with scars, as each season’s growth is very short and marked by two crescentic leaf-scars in addition to the ring-like scars left by the fall of the scales of the bud of the previous spring. The long shoots show similar ring-like scars at the base, and bear axillary buds


1 In some of the shrubby species the male catkins are solitary on the ends of the short shoots, and remain enclosed in the buds during winter, appearing in spring.

2 In B. nigra the fruit ripens in May or June.

3 The cones, scales, and fruits shown in Plates 269 and 270, were all drawn by Miss F.H. Woolward, except in the case of Figs. 8 and 16.

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