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The Trees of Great Britain and Ireland

nerves, with conspicuous tufts of rusty-brown hairs in the axils; petioles, glabrous or pubescent, 1½ to 1 inch long; stipules conspicuous, deciduous, ovate to lanceolate, obtuse, fringed with glandular hairs. The leaves turn blackish in autumn.

Flowers appearing very early, before the leaves, in February or March. Catkins, three to six in a raceme, at the tip of a branchlet. Staminate catkins, 2 to 4 inches long, at first erect and rigid, afterwards lax and pendent ; stamens,’ four, yellow, opposite the segments of the four-lobed calyx. Pistillate catkins, always erect, at first about ¼ inch long, smooth, with reddish-brown stigmas ; afterwards ½ inch long, ovoid, cone-like, the scales ending in purple shield-like expansions, each with a central brown point. Cones at first green, ultimately black, persistent on the tree after the escape of the nutlets. Nutlets obovate, blunt-angled, wingless or with a very narrow coriaceous wing.

The nutlets? are gradually shaken out of the cones by the wind during autumn and winter. Their walls are provided with small air-tight cavities, which enable them to float in water, and secrete an oil, which protects them from being wetted. Usually falling into streams and ditches, they float undamaged and unchanged during winter, and germinate in the water in early spring. The young seedlings, drifting to the bank, establish themselves where they happen to be stranded in a suitable place.

In winter, the twigs are glabrous and usually covered with a thin waxy secretion. Leaf-scars pentagonal or rhomboid, parallel to the twig on a projecting cushion, five- dotted, the lowermost three dots coalesced together. Stipule-scars linear, one on each side of a leaf-scar. Terminal bud present, similar to the lateral buds; all conspicuously stalked, ovoid, obtuse, with two external scales, viscid-glandular, and often covered with a purplish bloom. Pith triangular in section.

The common alder coppices freely from the stool; but rarely if ever produces root-suckers.

Varieties

The common alder, distributed over a wide area, shows considerable variation in the wild state, and several varieties have been described.

1. Var. barbata, Ledebour, Fl Rossica, iii. 6 57 (1851); Winkler, Betulaceæ, 118 (1904). Alnus barbata, C.A. Meyer, Verz. Pfl. Kauk. 43 (1831).

This variety is remarkably distinct in the foliage, but has the flowers and fruit of typical 4. glutinosa. Young branchlets pubescent. Leaves, about 3 inches long and 2 inches wide, rounded at the base, rounded or occasionally acute at the apex, margin with serrate lobules, ciliate; nerves eight or nine pairs, running parallel and curved to the margin; upper surface dark green, shining, minutely pubescent; lower surface pubescent, the pubescence densest along the midrib and nerves ; petiole ¾ inch, pubescent. Buds pubescent.

1 Kerner, in Nat. Hist. Plants, Eng. trans. ii, pp. 119, 133, 135, gives an elaborate account of the way in which pollination is effected by the wind, and of the devices for the protection of the pollen in rainy weather.

2 Cf. Miall’s account of the fruit of the alder in Round The Year, p. 279.